A foraging honeybee faces two problems: finding its way home from an unfamiliar patch of flowers, and telling her hive-mates exactly where to go. This lab shows both. A scout flies a crooked, wind-buffeted outbound path to a food source, continuously integrating every turn and every leg of flight into a running home vector — a process called path integration or dead reckoning. She then flies straight home along that vector, no matter how twisted the outward journey was. Back in the dark hive she performs the famous waggle dance, transposing the sun's compass bearing onto gravity so that other bees can decode direction and distance purely by touch, then fly out along the same route themselves.
Karl von Frisch won the 1973 Nobel Prize in Physiology or Medicine for decoding the waggle dance — one of the only confirmed cases of symbolic, referential communication known outside human language. Path integration was later confirmed with radar-tracked bees by Riley et al. (2005), who showed recruits fly almost exactly the vector encoded in the dance before switching to local search.
Watch a scout bee fly a wind-buffeted, crooked path to a food source while continuously integrating a home vector, fly straight home along that vector by dead reckoning, then perform a waggle dance that transposes the sun's angle onto gravity so recruited sisters can fly the same route.
Path integration lets a bee find home from any crooked outbound route; the sun compass supplies the reference bearing; the waggle dance re-encodes that bearing (relative to the sun) and the distance as a run angle and duration on the vertical comb.
Set the food source's compass angle and distance and the sun's azimuth, then watch the scout's crooked outbound trail, her straight dead-reckoned return, and the resulting dance. Toggle the vectors off to see just the bees, or speed the cycle up.
Karl von Frisch won the 1973 Nobel Prize in Physiology or Medicine for decoding the waggle dance — one of the few confirmed examples of symbolic, referential communication outside human language.